About the Authors

Karina Roxana Gheorghe

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institute, Karolinska University Hospital Solna, Stockholm, Sweden

Rogier M. Thurlings

Affiliation Division of Clinical Immunology and Rheumatology, Academic Medical Centre, University of Amsterdam, Amsterdam, The Netherlands

Marie Westman

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institute, Karolinska University Hospital Solna, Stockholm, Sweden

Maartje J. Boumans

Affiliation Division of Clinical Immunology and Rheumatology, Academic Medical Centre, University of Amsterdam, Amsterdam, The Netherlands

Vivianne Malmström

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institute, Karolinska University Hospital Solna, Stockholm, Sweden

Christina Trollmo

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institute, Karolinska University Hospital Solna, Stockholm, Sweden

Marina Korotkova

Affiliations Rheumatology Unit, Department of Medicine, Karolinska Institute, Karolinska University Hospital Solna, Stockholm, Sweden, Actar AB Inc., Stockholm, Sweden

Per-Johan Jakobsson

per-johan.jakobsson@ki.se

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institute, Karolinska University Hospital Solna, Stockholm, Sweden

Paul-Peter Tak

Affiliation Division of Clinical Immunology and Rheumatology, Academic Medical Centre, University of Amsterdam, Amsterdam, The Netherlands

Competing Interests

The authors have read the journal's policy and have the following conflicts: Profesor Tak has served as a consultant for Roche. Associate Professor Per-Johan Jakobsson is a member of the board of directors of NovaSAID. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: PJJ PPT. Performed the experiments: KRG RMT MW MJB. Analyzed the data: KRG MW MJB MK VM CT PJJ PPT. Contributed reagents/materials/analysis tools: RMT. Wrote the paper: KRG RMT MW MJB VM CT MK PJJ PPT.