About the Authors

Welkin H. Pope

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Deborah Jacobs-Sera

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Daniel A. Russell

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Craig L. Peebles

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Zein Al-Atrache

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Turi A. Alcoser

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Lisa M. Alexander

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Matthew B. Alfano

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Samantha T. Alford

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Nichols E. Amy

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Marie D. Anderson

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Alexander G. Anderson

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Andrew A. S. Ang

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Manuel Ares Jr.

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Amanda J. Barber

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Lucia P. Barker

Affiliation Howard Hughes Medical Institute, Science Education Alliance, Chevy Chase, Maryland United States of America

Jonathan M. Barrett

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

William D. Barshop

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Cynthia M. Bauerle

Address Current address: Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Ian M. Bayles

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Katherine L. Belfield

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Aaron A. Best

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Agustin Borjon Jr.

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Charles A. Bowman

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Christine A. Boyer

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Kevin W. Bradley

Affiliation Howard Hughes Medical Institute, Science Education Alliance, Chevy Chase, Maryland United States of America

Victoria A. Bradley

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Lauren N. Broadway

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Keshav Budwal

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Kayla N. Busby

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Ian W. Campbell

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Anne M. Campbell

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Alyssa Carey

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Steven M. Caruso

Affiliation Department of Biological Sciences, University of Maryland, Baltimore, Maryland, United States of America

Rebekah D. Chew

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Chelsea L. Cockburn

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Lianne B. Cohen

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Jeffrey M. Corajod

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Steven G. Cresawn

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Kimberly R. Davis

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Lisa Deng

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Dee R. Denver

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Breyon R. Dixon

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Sahrish Ekram

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Sarah C. R. Elgin

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Angela E. Engelsen

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Belle E. V. English

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Marcella L. Erb

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Crystal Estrada

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Laura Z. Filliger

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Ann M. Findley

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Lauren Forbes

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Mark H. Forsyth

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Tyler M. Fox

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Melissa J. Fritz

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Roberto Garcia

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Zindzi D. George

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Anne E. Georges

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Christopher R. Gissendanner

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Shannon Goff

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Rebecca Goldstein

Affiliation Department of Biological Sciences, University of Maryland, Baltimore, Maryland, United States of America

Kobie C. Gordon

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Russell D. Green

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Stephanie L. Guerra

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Krysta R. Guiney-Olsen

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Bridget G. Guiza

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Leila Haghighat

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Garrett V. Hagopian

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Catherine J. Harmon

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Jeremy S. Harmson

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Grant A. Hartzog

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Samuel E. Harvey

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Siping He

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Kevin J. He

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Kaitlin E. Healy

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Ellen R. Higinbotham

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Erin N. Hildebrandt

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Jason H. Ho

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Gina M. Hogan

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Victoria G. Hohenstein

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Nathan A. Holz

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Vincent J. Huang

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Ericka L. Hufford

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Peter M. Hynes

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Arrykka S. Jackson

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Erica C. Jansen

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Jonathan Jarvik

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Paul G. Jasinto

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Tuajuanda C. Jordan

Affiliation Howard Hughes Medical Institute, Science Education Alliance, Chevy Chase, Maryland United States of America

Tomas Kasza

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Murray A. Katelyn

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Jessica S. Kelsey

Affiliation Department of Biological Sciences, University of Maryland, Baltimore, Maryland, United States of America

Larisa A. Kerrigan

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Daryl Khaw

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Junghee Kim

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Justin Z. Knutter

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Ching-Chung Ko

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Gail V. Larkin

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Jennifer R. Laroche

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Asma Latif

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Kohana D. Leuba

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Sequoia I. Leuba

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Lynn O. Lewis

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Kathryn E. Loesser-Casey

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Courtney A. Long

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

A. Javier Lopez

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Nicholas Lowery

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Tina Q. Lu

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Victor Mac

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Isaac R. Masters

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Jazmyn J. McCloud

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Molly J. McDonough

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Andrew J. Medenbach

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Anjali Menon

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Rachel Miller

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Brandon K. Morgan

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Patrick C. Ng

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Elvis Nguyen

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Katrina T. Nguyen

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Emilie T. Nguyen

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Kaylee M. Nicholson

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Lindsay A. Parnell

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Caitlin E. Peirce

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Allison M. Perz

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Luke J. Peterson

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Rachel E. Pferdehirt

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Seegren V. Philip

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Kit Pogliano

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Joe Pogliano

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Tamsen Polley

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Erica J. Puopolo

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Hannah S. Rabinowitz

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Michael J. Resiss

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Corwin N. Rhyan

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Yetta M. Robinson

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Lauren L. Rodriguez

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Andrew C. Rose

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Jeffrey D. Rubin

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Jessica A. Ruby

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Margaret S. Saha

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

James W. Sandoz

Affiliation Department of Biological Sciences, University of Maryland, Baltimore, Maryland, United States of America

Judith Savitskaya

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Dale J. Schipper

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Christine E. Schnitzler

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Amanda R. Schott

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

J. Bradley Segal

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Christopher D. Shaffer

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Kathryn E. Sheldon

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Erica M. Shepard

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Jonathan W. Shepardson

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Madav K. Shroff

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Jessica M. Simmons

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Erika F. Simms

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Brandy M. Simpson

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Kathryn M. Sinclair

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Robert L. Sjoholm

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Ingrid J. Slette

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Blaire C. Spaulding

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Clark L. Straub

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Joseph Stukey

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Trevor Sughrue

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

Tin-Yun Tang

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Lyons M. Tatyana

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Stephen B. Taylor

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Barbara J. Taylor

Affiliation Department of Zoology, Oregon State University, Corvallis, Oregon, United States of America

Louise M. Temple

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Jasper V. Thompson

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Michael P. Tokarz

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Stephanie E. Trapani

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Alexander P. Troum

Affiliation Department of Biology, James Madison University, Harrisonburg, Virginia, United States of America

Jonathan Tsay

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Anthony T. Tubbs

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Jillian M. Walton

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Danielle H. Wang

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Hannah Wang

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

John R. Warner

Affiliation Department of Biology, University of Louisiana at Monroe, Monroe, Louisiana, United States of America

Emilie G. Weisser

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Samantha C. Wendler

Affiliation Department of Biological Sciences, University of Mary Washington, Fredericksburg, Virginia, United States of America

Kathleen A. Weston-Hafer

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Hilary M. Whelan

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Kurt E. Williamson

Affiliation Biology Department, College of William & Mary, Williamsburg, Virginia, United States of America

Angelica N. Willis

Affiliation Biology Department, A. Paul Schaap Science Center, Hope College, Holland, Michigan, United States of America

Hannah S. Wirtshafter

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Theresa W. Wong

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Phillip Wu

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Yun jeong Yang

Affiliation Division of Biological Sciences, University of California San Diego, La Jolla, California, United States of America

Brandon C. Yee

Affiliation Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America

David A. Zaidins

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Bo Zhang

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Melina Y. Zúniga

Affiliation Department of Biology, Spelman College, Atlanta, Georgia, United States of America

Roger W. Hendrix

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Graham F. Hatfull

gfh@pitt.edu

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

For the list of Author Contributions please see Table S1.