About the Authors

Giulia Pinton

Affiliation Department of Chemical, Food, Pharmaceutical and Pharmacological Sciences, Drug and Food Biotechnology Center, University of Piemonte Orientale A. Avogadro, Novara, Italy

Warren Thomas

Affiliation Department of Molecular Medicine, Royal College of Surgeons in Ireland and Education and Research Centre, Beaumont Hospital, Dublin, Ireland

Paolo Bellini

Affiliation Department of Chemical, Food, Pharmaceutical and Pharmacological Sciences, Drug and Food Biotechnology Center, University of Piemonte Orientale A. Avogadro, Novara, Italy

Arcangela Gabriella Manente

Affiliation Department of Chemical, Food, Pharmaceutical and Pharmacological Sciences, Drug and Food Biotechnology Center, University of Piemonte Orientale A. Avogadro, Novara, Italy

Roberto E. Favoni

Affiliation Laboratory of Experimental Pharmacology, National Cancer Institute, Genoa, Italy

Brian J. Harvey

Affiliation Department of Molecular Medicine, Royal College of Surgeons in Ireland and Education and Research Centre, Beaumont Hospital, Dublin, Ireland

Luciano Mutti

Contributed equally to this work with: Luciano Mutti, Laura Moro

Affiliation Department of Medicine, Local Health Unit 11, Vercelli, Italy

Laura Moro

Contributed equally to this work with: Luciano Mutti, Laura Moro

moro@pharm.unipmn.it

Affiliation Department of Chemical, Food, Pharmaceutical and Pharmacological Sciences, Drug and Food Biotechnology Center, University of Piemonte Orientale A. Avogadro, Novara, Italy

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GP WT L. Moro. Performed the experiments: GP WT PB AGM. Analyzed the data: GP WT REF BH L. Mutti, L. Moro. Contributed reagents/materials/analysis tools: REF BH L. Mutti, L. Moro. Wrote the paper: BH L. Mutti, L. Moro.