About the Authors

Alessandro Biffi

Affiliations Center for Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Hemorrhagic Stroke Research Group, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Anna Plourde

Affiliations Center for Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Hemorrhagic Stroke Research Group, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Yiping Shen

Affiliation Center for Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America

Robert Onofrio

Affiliation Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Eric E. Smith

Affiliation Department of Clinical Neurosciences, University of Calgary, Calgary, Alberta, Canada

Matthew Frosch

Affiliation Massachusetts General Hospital Institute for Neurodegenerative Disease, Massachusetts General Hospital, Boston, Massachusetts, United States of America

Claudia M. Prada

Affiliation Massachusetts General Hospital Institute for Neurodegenerative Disease, Massachusetts General Hospital, Boston, Massachusetts, United States of America

James Gusella

Affiliations Center for Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Steven M. Greenberg

Affiliation Hemorrhagic Stroke Research Group, Massachusetts General Hospital, Boston, Massachusetts, United States of America

Jonathan Rosand

jrosand@partners.org

Affiliations Center for Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Hemorrhagic Stroke Research Group, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YS EES MPF CMP JFG SMG JR. Performed the experiments: AP YS RO. Analyzed the data: AB RO MPF. Contributed reagents/materials/analysis tools: JR. Wrote the paper: AB AP YS EES MPF CMP JFG SMG JR.