About the Authors

Thomas J. Hoffmann

Affiliation Department of Epidemiology and Biostatistics and Institute of Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Nicholas J. Marini

Affiliation Department of Molecular and Cellular Biology, California Institute for Quantitative Biosciences, University of California, Berkeley, California, United States of America

John S. Witte

WitteJ@humgen.ucsf.edu

Affiliation Department of Epidemiology and Biostatistics and Institute of Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TJH NJM JSW. Performed the experiments: TJH. Wrote the paper: TJH NJM JSW.