About the Authors

Thomas W. Lewis

tom.lewis@aht.org.uk

Affiliation Kennel Club Genetics Centre at the Animal Health Trust, Animal Health Trust, Newmarket, United Kingdom

Sarah C. Blott

Affiliation Kennel Club Genetics Centre at the Animal Health Trust, Animal Health Trust, Newmarket, United Kingdom

John A. Woolliams

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, United Kingdom

Competing Interests

Hip score and pedigree data was collated and provided by the UK Kennel Club.

Author Contributions

Conceived and designed the experiments: TWL SCB JAW. Performed the experiments: TWL. Analyzed the data: TWL JAW. Wrote the paper: TWL JAW.