About the Authors

Ricardo Betancur-R

betanri@gmail.com

Address Current address: Department of Biological Sciences, The George Washington University, Washington, D. C., United States of America

Affiliations Department of Biological Sciences, Auburn University, Auburn, Alabama, United States of America, Universidad Nacional de Colombia sede Caribe, CECIMAR/INVEMAR, Cerro Punta Betín, Santa Marta, Colombia

Arturo Acero P.

Affiliation Universidad Nacional de Colombia sede Caribe, CECIMAR/INVEMAR, Cerro Punta Betín, Santa Marta, Colombia

Hermann Duque-Caro

Affiliation Duque Caro y Cia. Ltda., Bogotá, Colombia

Scott R. Santos

Affiliation Department of Biological Sciences, Auburn University, Auburn, Alabama, United States of America

Competing Interests

The only role of Duque Caro y Cia. Ltda. in the study was to support the salary of the third author (H. Duque-Caro). This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RBR AAP SRS. Performed the experiments: RBR. Analyzed the data: RBR HDC SRS. Contributed reagents/materials/analysis tools: RBR. Wrote the paper: RBR AAP HDC SRS.