About the Authors

Kevin E. Doherty


Affiliation National Audubon Society Science, Wyoming Audubon, Laramie, Wyoming, United States of America

David E. Naugle

Affiliation Wildlife Biology Program, University of Montana, Missoula, Montana, United States of America

Jeffrey S. Evans

Affiliation North America Science, The Nature Conservancy, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KED DN. Performed the experiments: KED. Analyzed the data: KED JSE. Wrote the paper: KED DN.