About the Authors

Colin P. Groves


Affiliation School of Archaeology & Anthropology, Australian National University, Canberra, Australian Capital Territory, Australia

Prithiviraj Fernando

Affiliation Centre for Conservation and Research, Rajagiriya, Sri Lanka

Jan Robovský

Affiliation Department of Zoology, Faculty of Science, University of South Bohemia, České Budějovice, Czech Republic

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PF CPG. Performed the experiments: PF. Analyzed the data: PF CPG JR. Contributed reagents/materials/analysis tools: PF JR. Wrote the paper: PF CPG JR.