About the Authors

Guoliang Li

Affiliation Computational & Mathematical Biology, Genome Institute of Singapore, Singapore, Singapore

Jian Ma

Affiliation Department of Bioengineering, University of Illinois at Urbana-Champaign, Urbana, Illinois, United States of America

Louxin Zhang

matzlx@nus.edu.sg

Affiliation Department of Mathematics, National University of Singapore, Singapore, Singapore

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GL LZ. Performed the experiments: GL JM. Analyzed the data: GL JM. Wrote the paper: JM LZ.