About the Authors

Asako Shindo

Affiliation Department of Developmental Biology, National Institute for Basic Biology, Okazaki, Aichi, Japan

Yusuke Hara

Affiliations Department of Developmental Biology, National Institute for Basic Biology, Okazaki, Aichi, Japan, Department of Basic Biology, School of Life Science, The Graduate University of Advanced Studies (SOKENDAI), Okazaki, Aichi, Japan

Takamasa S. Yamamoto

Affiliation Department of Developmental Biology, National Institute for Basic Biology, Okazaki, Aichi, Japan

Masamichi Ohkura

Affiliation Saitama University Brain Science Institute, Saitama, Saitama, Japan

Junichi Nakai

Affiliation Saitama University Brain Science Institute, Saitama, Saitama, Japan

Naoto Ueno

ueno@nibb.ac.jp

Affiliations Department of Developmental Biology, National Institute for Basic Biology, Okazaki, Aichi, Japan, Department of Basic Biology, School of Life Science, The Graduate University of Advanced Studies (SOKENDAI), Okazaki, Aichi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AS NU. Performed the experiments: AS YH TSY. Analyzed the data: AS. Contributed reagents/materials/analysis tools: AS MO JN. Wrote the paper: AS NU.