About the Authors

Adam O. Whelan

Affiliation TB Research Group, Veterinary Laboratories Agency Weybridge, New Haw, United Kingdom

Michael Coad

Affiliation TB Research Group, Veterinary Laboratories Agency Weybridge, New Haw, United Kingdom

Paul J. Cockle

Current address: Vaccines Research, Pfizer Global Research and Development, Sandwich, United Kingdom

Affiliation TB Research Group, Veterinary Laboratories Agency Weybridge, New Haw, United Kingdom

Glyn Hewinson

Affiliation TB Research Group, Veterinary Laboratories Agency Weybridge, New Haw, United Kingdom

Martin Vordermeier

Affiliation TB Research Group, Veterinary Laboratories Agency Weybridge, New Haw, United Kingdom

Stephen V. Gordon

stephen.gordon@ucd.ie

Affiliations TB Research Group, Veterinary Laboratories Agency Weybridge, New Haw, United Kingdom, School of Agriculture, Food Science and Veterinary Medicine, University College Dublin, Dublin, Ireland, School of Medicine and Medical Science, University College Dublin, Dublin, Ireland, School of Biomolecular and Biomedical Science, College of Life Sciences, University College Dublin, Dublin, Ireland, Conway Institute of Biomolecular and Biomedical Research, University College Dublin, Dublin, Ireland

Competing Interests

While PJC now works for Pfizer, the work in this manuscript was performed while PJC was a Veterinary Laboratories Agency employee, and Pfizer had no role as a funder of this work.

Author Contributions

Conceived and designed the experiments: GH MV SVG. Performed the experiments: AW MC PJC MV SVG. Analyzed the data: AW PJC GH MV SVG. Wrote the paper: AW MV SVG.