About the Authors

Karim Ouattara

Affiliations Laboratoire EthoS “Ethologie animale et humaine”, U.M.R. 6552-C.N.R.S., Université de Rennes 1, Station Biologique, Paimpont, France, Centre Suisse de Recherches Scientifiques, Abidjan, Côte d'Ivoire, Laboratoire de Zoologie et de Biologie Animale, Université de Cocody, Abidjan, Côte d'Ivoire

Alban Lemasson

Affiliation Laboratoire EthoS “Ethologie animale et humaine”, U.M.R. 6552-C.N.R.S., Université de Rennes 1, Station Biologique, Paimpont, France

Klaus Zuberbühler

kz3@st-and.ac.uk

Affiliations Centre Suisse de Recherches Scientifiques, Abidjan, Côte d'Ivoire, School of Psychology, University of St Andrews, St Andrews, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KO AL KZ. Performed the experiments: KO. Analyzed the data: KO. Contributed reagents/materials/analysis tools: AL KZ. Wrote the paper: AL KZ.