About the Authors

Hanna Göransson

Affiliation Department of Medical Sciences, Uppsala University, Uppsala, Sweden

Karolina Edlund

Affiliation Department of Genetics and Pathology, Uppsala University, Uppsala, Sweden

Maria Rydåker

Affiliation Department of Medical Sciences, Uppsala University, Uppsala, Sweden

Markus Rasmussen

Affiliation Department of Medical Sciences, Uppsala University, Uppsala, Sweden

Johan Winquist

Affiliation Department of Medical Sciences, Uppsala University, Uppsala, Sweden

Simon Ekman

Affiliation Department of Oncology, Uppsala University Hospital, Uppsala, Sweden

Michael Bergqvist

Affiliation Department of Oncology, Uppsala University Hospital, Uppsala, Sweden

Andrew Thomas

Affiliation AstraZeneca, Alderley Park, Macclesfield, United Kingdom

Mats Lambe

Affiliations Department of Medical Epidemiology and Biostatistics, Karolinska Institute, Stockholm, Sweden, Regional Oncologic Centre, Uppsala University Hospital, Uppsala, Sweden

Richard Rosenquist

Affiliation Department of Genetics and Pathology, Uppsala University, Uppsala, Sweden

Lars Holmberg

Affiliations Regional Oncologic Centre, Uppsala University Hospital, Uppsala, Sweden, Division of Cancer Studies, Medical School, King's College London, London, United Kingdom, Department of Surgical Sciences, Uppsala University, Uppsala, Sweden

Patrick Micke

Affiliation Department of Genetics and Pathology, Uppsala University, Uppsala, Sweden

Johan Botling

Affiliation Department of Genetics and Pathology, Uppsala University, Uppsala, Sweden

Anders Isaksson

anders.isaksson@medsci.uu.se

Affiliation Department of Medical Sciences, Uppsala University, Uppsala, Sweden

Competing Interests

The co-author Andrew P. Thomas is employed by AstraZeneca. Anders Isaksson and Andrew P. Thomas hold stock in Astra Zeneca.

Author Contributions

Conceived and designed the experiments: HG AI. Performed the experiments: KE MR JB. Analyzed the data: HG MR JW JB AI. Contributed reagents/materials/analysis tools: SE MB APT ML RR LH PM JB AI. Wrote the paper: HG KE AI.