About the Authors

Scott H. Newman

scott.newman@fao.org

Affiliations EMPRES Wildlife Unit, Infectious Disease Group, Emergency Centre for Transboundary Animal Diseases, Animal Production and Health Division, Food & Agriculture Organization of the United Nations, Rome, Italy, Wildlife Conservation Society, Field Veterinary Program, Bronx, New York, United States of America

Samuel A. Iverson

Affiliation United States Geological Survey, Western Ecological Research Center, Vallejo, California, United States of America

John Y. Takekawa

Affiliation United States Geological Survey, Western Ecological Research Center, Vallejo, California, United States of America

Martin Gilbert

Affiliation Wildlife Conservation Society, Field Veterinary Program, Bronx, New York, United States of America

Diann J. Prosser

Affiliation United States Geological Survey, Patuxent Wildlife Research Center, Beltsville, Maryland, United States of America

Nyambyar Batbayar

Affiliation Wildlife Science Conservation Center of Mongolia, and Ornithological Laboratory, Institute of Biology, Mongolia Academy of Sciences, Ulaanbaatar, Mongolia

Tseveenmyadag Natsagdorj

Affiliation Ornithological Laboratory, Institute of Biology, Mongolia Academy of Sciences, Ulaanbaatar, Mongolia

David C. Douglas

Affiliation United States Geological Survey, Alaska Science Centre, Juneau, Alaska, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SHN JYT. Performed the experiments: SHN JYT MG DJP NB TN. Analyzed the data: SAI DCD. Contributed reagents/materials/analysis tools: SHN JYT MG DJP NB TN DCD. Wrote the paper: SHN SAI.