About the Authors

Shinya Watanabe

Current address: Tuberculosis Research Section, Laboratory of Clinical Infectious Disease, National Institute of Allergy and Infectious Disease, National Institutes of Health, Bethesda, Maryland, United States of America

Affiliation Department of Infection Control Science, Graduate School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan

Teruyo Ito

teruybac@juntendo.ac.jp

Affiliations Department of Infection Control Science, Graduate School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan, Department of Bacteriology, School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan

Takashi Sasaki

Affiliation Department of Infection Control Science, Graduate School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan

Shanshuang Li

Affiliation Department of Infection Control Science, Graduate School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan

Ikuo Uchiyama

Affiliation National Institute for Basic Biology, National Institutes of Natural Sciences, Myodaiji, Okazaki, Japan

Kozue Kishii

Current address: Laboratory of Molecular Epidemiology for Infectious Agents, Kitasato Institute for Life Sciences, Kitasato University, Minato-ku, Tokyo, Japan

Affiliation Department of Infection Control Science, Graduate School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan

Ken Kikuchi

Affiliations Department of Infection Control Science, Graduate School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan, Department of Bacteriology, School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan

Robert Leo Skov

Affiliation National Center for Antimicrobials and Infection Control, Statens Serum Institut, Copenhagen, Denmark

Keiichi Hiramatsu

Affiliations Department of Infection Control Science, Graduate School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan, Department of Bacteriology, School of Medicine, Juntendo University, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SW TI KH. Performed the experiments: SW TI TS SL KK. Analyzed the data: SW TI IU KH. Contributed reagents/materials/analysis tools: TS KK KK RS. Wrote the paper: SW TI RS KH.