About the Authors

Naohito Kitamura

Contributed equally to this work with: Naohito Kitamura, Runa Araya

Affiliation Yamada Research Unit, RIKEN Brain Science Institute, Saitama, Japan

Runa Araya

Contributed equally to this work with: Naohito Kitamura, Runa Araya

Affiliation Yamada Research Unit, RIKEN Brain Science Institute, Saitama, Japan

Moeko Kudoh

Affiliation Yamada Research Unit, RIKEN Brain Science Institute, Saitama, Japan

Haruo Kishida

Affiliation Yamada Research Unit, RIKEN Brain Science Institute, Saitama, Japan

Tetsuya Kimura

Affiliation Laboratory for Alzheimer's Diseases, RIKEN Brain Science Institute, Saitama, Japan

Miyuki Murayama

Affiliation Laboratory for Alzheimer's Diseases, RIKEN Brain Science Institute, Saitama, Japan

Akihiko Takashima

Affiliation Laboratory for Alzheimer's Diseases, RIKEN Brain Science Institute, Saitama, Japan

Yuriko Sakamaki

Affiliation Research Resource Center, RIKEN Brain Science Institute, Saitama, Japan

Tsutomu Hashikawa

Affiliation Research Resource Center, RIKEN Brain Science Institute, Saitama, Japan

Shingo Ito

Affiliation Department of Molecular Biopharmacy and Genetics, Tohoku University, Sendai, Japan

Sumio Ohtsuki

Affiliation Department of Molecular Biopharmacy and Genetics, Tohoku University, Sendai, Japan

Tetsuya Terasaki

Affiliation Department of Molecular Biopharmacy and Genetics, Tohoku University, Sendai, Japan

Jürgen Wess

Affiliation Laboratory of Bioorganic Chemistry, National Institute of Diabetes and Digestive and Kidney Diseases, Bethesda, Maryland, United States of America

Masahisa Yamada

masahisa@brain.riken.jp

Affiliation Yamada Research Unit, RIKEN Brain Science Institute, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MY. Performed the experiments: NK RA MK HK TK MM AT YS SI SO. Analyzed the data: TK TH JW. Contributed reagents/materials/analysis tools: TT. Wrote the paper: JW MY.