About the Authors

Elizabeth A. Gaskell

Affiliation Institute of Integrative and Comparative Biology, University of Leeds, Leeds, United Kingdom

Judith E. Smith

Affiliation Institute of Integrative and Comparative Biology, University of Leeds, Leeds, United Kingdom

John W. Pinney

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Dave R. Westhead

Affiliation Institute of Molecular and Cellular Biology, University of Leeds, Leeds, United Kingdom

Glenn A. McConkey

G.A.McConkey@Leeds.ac.uk

Affiliation Institute of Integrative and Comparative Biology, University of Leeds, Leeds, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EAG GAM. Performed the experiments: EAG GAM. Analyzed the data: EAG JWP DRW GAM. Contributed reagents/materials/analysis tools: JS JWP. Wrote the paper: EAG JWP DRW GAM.