About the Authors

Michelle V. Wagner

Affiliations Howard Hughes Medical Institute, University of California San Diego School of Medicine, La Jolla, California, United States of America, Department of Cellular & Molecular Medicine, University of California San Diego School of Medicine, La Jolla, California, United States of America, Biomedical Science Graduate Program, University of California San Diego School of Medicine, La Jolla, California, United States of America

Marcus B. Smolka

Affiliations Department of Cellular & Molecular Medicine, University of California San Diego School of Medicine, La Jolla, California, United States of America, Ludwig Institute for Cancer Research, University of California San Diego School of Medicine, La Jolla, California, United States of America

Rob A. M. de Bruin

Affiliation Department of Molecular Biology, The Scripps Research Institute, La Jolla, California, United States of America

Huilin Zhou

Affiliations Department of Cellular & Molecular Medicine, University of California San Diego School of Medicine, La Jolla, California, United States of America, Ludwig Institute for Cancer Research, University of California San Diego School of Medicine, La Jolla, California, United States of America

Curt Wittenberg

Affiliations Department of Molecular Biology, The Scripps Research Institute, La Jolla, California, United States of America, Department of Cell Biology, The Scripps Research Institute, La Jolla, California, United States of America

Steven F. Dowdy

sdowdy@ucsd.edu

Affiliations Howard Hughes Medical Institute, University of California San Diego School of Medicine, La Jolla, California, United States of America, Department of Cellular & Molecular Medicine, University of California San Diego School of Medicine, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MVW SFD. Performed the experiments: MVW MBS RAMdB. Analyzed the data: MVW MBS RAMdB HZ CW. Contributed reagents/materials/analysis tools: RAMdB HZ CW. Wrote the paper: MVW SFD.