About the Authors

Zheng Wang

wang.zheng@yale.edu

Affiliation Department of Ecology and Evolutionary Biology, Yale University, New Haven, Connecticut, United States of America

Peter R. Johnston

Affiliation Herbarium PDD, Landcare Research, Auckland, New Zealand

Zhu L. Yang

Affiliation Key Laboratory of Biodiversity and Biogeography, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, China

Jeffrey P. Townsend

Affiliation Department of Ecology and Evolutionary Biology, Yale University, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZW JPT. Performed the experiments: ZW. Analyzed the data: ZW JPT. Contributed reagents/materials/analysis tools: PRJ ZY. Wrote the paper: ZW PRJ ZY JPT.