About the Authors

John F. Heidelberg

jheidelb@usc.edu

Affiliation Department of Biological Sciences, Marine Environmental Biology Division, Wrigley Institute for Environmental Studies, University of Southern California, Avalon, California, United States of America

William C. Nelson

Affiliation Department of Biological Sciences, Marine Environmental Biology Division, Wrigley Institute for Environmental Studies, University of Southern California, Avalon, California, United States of America

Thomas Schoenfeld

Affiliation Lucigen Corporation, Middleton, Wisconsin, United States of America

Devaki Bhaya

Affiliation Department of Plant Biology, Carnegie Institution for Science, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JFH WCN DB. Performed the experiments: JFH DB. Analyzed the data: JFH WCN TS DB. Wrote the paper: JFH WCN TS DB.