About the Authors

Lee Smith

l.smith@hrsu.mrc.ac.uk

Current address: MRC Human Reproductive Sciences Unit, The Queen's Medical Research Institute, Edinburgh, United Kingdom

Affiliation Mammalian Genetics Unit, MRC Harwell, Oxfordshire, United Kingdom

John Willan

Affiliation Mammalian Genetics Unit, MRC Harwell, Oxfordshire, United Kingdom

Nick Warr

Affiliation Mammalian Genetics Unit, MRC Harwell, Oxfordshire, United Kingdom

Frances A. Brook

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Michael Cheeseman

Affiliation Mammalian Genetics Unit, MRC Harwell, Oxfordshire, United Kingdom

Richard Sharpe

Affiliation MRC Human Reproductive Sciences Unit, The Queen's Medical Research Institute, Edinburgh, United Kingdom

Pam Siggers

Affiliation Mammalian Genetics Unit, MRC Harwell, Oxfordshire, United Kingdom

Andy Greenfield

Affiliation Mammalian Genetics Unit, MRC Harwell, Oxfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LS PS AG. Performed the experiments: LS JW MC RS PS. Analyzed the data: LS MC RS PS AG. Contributed reagents/materials/analysis tools: NW FB. Wrote the paper: LS AG.