About the Authors

Yutaka Sakai

Affiliation Brain Science Institute, Tamagawa University, Machida, Tokyo, Japan

Tomoki Fukai


Affiliation Laboratory for Neural Circuit Theory, Brain Science Institute, RIKEN, Wako, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YS. Performed the experiments: YS. Wrote the paper: TF.