About the Authors

Naomi D. Willis

Affiliation School of Biological and Biomedical Sciences, Durham University, Durham, United Kingdom

Thomas R. Cox

Affiliation School of Biological and Biomedical Sciences, Durham University, Durham, United Kingdom

Syed F. Rahman-Casañs

Affiliation James Cook University Hospital, Middlesbrough, United Kingdom

Kim Smits

Affiliation Department of Epidemiology, University of Maastricht, Maastricht, The Netherlands

Stefan A. Przyborski

Affiliation School of Biological and Biomedical Sciences, Durham University, Durham, United Kingdom

Piet van den Brandt

Affiliation Department of Epidemiology, University of Maastricht, Maastricht, The Netherlands

Manon van Engeland

Affiliation Department of Pathology, University of Maastricht, Maastricht, The Netherlands

Matty Weijenberg

Affiliation Department of Epidemiology, University of Maastricht, Maastricht, The Netherlands

Robert G. Wilson

Affiliation James Cook University Hospital, Middlesbrough, United Kingdom

Adriaan de Bruïne

Affiliation Department of Pathology, University of Maastricht, Maastricht, The Netherlands

Christopher J. Hutchison

c.j.hutchison@durham.ac.uk

Affiliation School of Biological and Biomedical Sciences, Durham University, Durham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RW AdB CH. Performed the experiments: NDW TC SRC. Analyzed the data: NDW TC KS SP PAvdB MW MvE. Contributed reagents/materials/analysis tools: PAvdB MW MvE. Wrote the paper: CH.