About the Authors

Catriona M. Harris

catriona@mcs.st-and.ac.uk

Affiliations Sea Mammal Research Unit, University of St. Andrews, St Andrews, Fife, United Kingdom, Centre for Research into Ecological and Environmental Modelling, University of St Andrews, St Andrews, Fife, United Kingdom

Justin M. J. Travis

Affiliation School of Biological Sciences, University of Aberdeen, Aberdeen, Aberdeenshire, United Kingdom

John Harwood

Affiliations Sea Mammal Research Unit, University of St. Andrews, St Andrews, Fife, United Kingdom, Centre for Research into Ecological and Environmental Modelling, University of St Andrews, St Andrews, Fife, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CMH JMT JH. Performed the experiments: CMH. Analyzed the data: CMH. Contributed reagents/materials/analysis tools: JMT. Wrote the paper: CMH JH.