About the Authors

Jo N. Fleming

To whom correspondence should be addressed. E-mail: flemij@u.washington.edu

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

Richard A. Nash

Affiliation Clinical Research Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

D. O. McLeod

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

David F. Fiorentino

Affiliation Department of Dermatology, Stanford University School of Medicine, Stanford, California, United States of America

Howard M. Shulman

Affiliation Clinical Research Division, Pathology Section, Fred Hutchinson Cancer Research Center and Seattle Cancer Care Alliance, Seattle, Washington, United States of America

M. Kari Connolly

Affiliation Department of Medicine, University of California at San Francisco, San Francisco, California, United States of America

Jerry A. Molitor

Affiliation Department of Rheumatology, Virginia Mason Medical Center, Seattle, Washington, United States of America

Gretchen Henstorf

Affiliation Program in Transplantation Biology, Clinical Research Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Robert Lafyatis

Affiliation Rheumatology Section, Boston University School of Medicine, Boston, Massachusetts, United States of America

David K. Pritchard

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

Lawrence D. Adams

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

Daniel E. Furst

Affiliation Department of Rheumatology, David Geffen School of Medicine, University of California at Los Angeles, Los Angeles, California, United States of America

Stephen M. Schwartz

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS JF DF. Performed the experiments: JF DM. Analyzed the data: JF GH HS. Contributed reagents/materials/analysis tools: JF RN DF MC JM GH RL HS. Wrote the paper: SS JF RN JM RL DP LA DF HS.