About the Authors

Ari Ueno

To whom correspondence should be addressed. E-mail: aueno@ecs.c.u-tokyo.ac.jp

Affiliation The University of Tokyo, Tokyo, Japan

Satoshi Hirata

Affiliation Great Ape Research Institute, Hayashibara Biochemical Laboratories, Tamano, Okayama, Japan

Kohki Fuwa

Affiliation Great Ape Research Institute, Hayashibara Biochemical Laboratories, Tamano, Okayama, Japan

Keiko Sugama

Affiliation Great Ape Research Institute, Hayashibara Biochemical Laboratories, Tamano, Okayama, Japan

Kiyo Kusunoki

Affiliation Great Ape Research Institute, Hayashibara Biochemical Laboratories, Tamano, Okayama, Japan

Goh Matsuda

Affiliation The University of Tokyo, Tokyo, Japan

Hirokata Fukushima

Affiliation The University of Tokyo, Tokyo, Japan

Kazuo Hiraki

Affiliation The University of Tokyo, Tokyo, Japan

Masaki Tomonaga

Affiliation Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Toshikazu Hasegawa

Affiliation The University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MT AU GM KH TH SH. Performed the experiments: KF KS KK SH. Analyzed the data: AU HF. Contributed reagents/materials/analysis tools: GM. Wrote the paper: AU GM HF.