About the Authors

Ruby Hsu

Affiliation Department of Neuroscience, Columbia University, New York, New York, United States of America

Abigail Woodroffe

Affiliation Department of Epidemiology, Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Wen-Sung Lai

Affiliation Department of Physiology and Cellular Biophysics, College of Physicians and Surgeons, Columbia University, New York, New York, United States of America

Melloni N. Cook

Affiliation Department of Psychology, The University of Memphis, Memphis, Tennessee, United States of America

Jun Mukai

Affiliation Department of Physiology and Cellular Biophysics, College of Physicians and Surgeons, Columbia University, New York, New York, United States of America

Jonathan P. Dunning

Affiliation Department of Psychology, The University of Memphis, Memphis, Tennessee, United States of America

Douglas J. Swanson

Affiliation Department of Anatomy and Neurobiology, University of Tennessee Health Science Center, Memphis, Tennessee, United States of America

J. Louw Roos

Affiliation Department of Psychiatry, University of Pretoria and Weskoppies Hospital, Pretoria, Republic of South Africa

Gonçalo R. Abecasis

Affiliation Department of Biostatistics, Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Maria Karayiorgou

To whom correspondence should be addressed. E-mail: mk2758@columbia.edu (MK); jag90@columbia.edu (JG)

Affiliation Department of Psychiatry, College of Physicians and Surgeons, Columbia University, New York, New York, United States of America

Joseph A. Gogos

To whom correspondence should be addressed. E-mail: mk2758@columbia.edu (MK); jag90@columbia.edu (JG)

Affiliations Department of Neuroscience, Columbia University, New York, New York, United States of America, Department of Physiology and Cellular Biophysics, College of Physicians and Surgeons, Columbia University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JG MK. Performed the experiments: RH WL JM MC JD DS. Analyzed the data: GA JG MK RH WL MC JD DS AW. Contributed reagents/materials/analysis tools: JR. Wrote the paper: GA JG MK RH WL MC AW.