About the Authors

Caitlin Black

Contributed equally to this work with: Caitlin Black, Tom Hart

caitlin.black@lincoln.ox.ac.uk

Affiliation University of Oxford, Department of Zoology, Oxford, United Kingdom

Ben Collen

‡ These authors also contributed equally to this work.

Affiliation University College London, Centre for Biodiversity & Environment Research, London, United Kingdom

Daniel Johnston

‡ These authors also contributed equally to this work.

Affiliation British Antarctic Survey, King Edward Point Research Station, Cumberland East Bay, South Georgia

Tom Hart

Contributed equally to this work with: Caitlin Black, Tom Hart

Affiliation University of Oxford, Department of Zoology, Oxford, United Kingdom

Competing Interests

Funding was provided by Quark Expeditions Ltd., and Exodus Travel Ltd. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CB TH. Performed the experiments: CB TH. Analyzed the data: CB. Contributed reagents/materials/analysis tools: BC DJ TH. Wrote the paper: CB BC TH.