About the Authors

Will T. S. Miles

w.miles@abdn.ac.uk

Affiliation Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen, Aberdeenshire, United Kingdom

Roddy Mavor

Affiliation Joint Nature Conservation Committee, Aberdeen, Aberdeenshire, United Kingdom

Nick J. Riddiford

Affiliation Fair Isle Bird Observatory, Fair Isle, Shetland, United Kingdom

Paul V. Harvey

Affiliation Shetland Biological Records Centre, Shetland Amenity Trust, Garthspool, Lerwick, Shetland, United Kingdom

Roger Riddington

Affiliation Spindrift, Eastshore, Virkie, Shetland, United Kingdom

Deryk N. Shaw

Affiliation Fair Isle Bird Observatory, Fair Isle, Shetland, United Kingdom

David Parnaby

Affiliation Fair Isle Bird Observatory, Fair Isle, Shetland, United Kingdom

Jane M. Reid

Affiliation Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen, Aberdeenshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WTSM RM NJR PVH RR DNS DP JMR. Performed the experiments: WTSM RM NJR PVH RR DNS DP JMR. Analyzed the data: WTSM RM JMR. Contributed reagents/materials/analysis tools: WTSM RM JMR. Wrote the paper: WTSM RM NJR PVH RR DNS DP JMR. Carried out and supervised fieldwork: WTSM RM NJR PVH RR DNS DP JMR.