About the Authors

Chie Sugimoto

Contributed equally to this work with: Chie Sugimoto, Hiroshi Wakao

Affiliation Department of Hygiene & Cellular Preventive Medicine, Graduate School of Medicine, Hokkaido University, Sapporo, 060–8638, Japan

Takahiko Konno

Affiliation Department of Rheumatology, Tokeidai-Memorial Clinic, Sapporo, 060–0031, Japan

Rika Wakao

Affiliation Pharmaceutical and Medical Device Agency (PMDA), Kasumigaseki, Tokyo, 100–0013, Japan

Hiroko Fujita

Affiliation Department of Hygiene & Cellular Preventive Medicine, Graduate School of Medicine, Hokkaido University, Sapporo, 060–8638, Japan

Hiroyoshi Fujita

Affiliation Department of Hygiene & Cellular Preventive Medicine, Graduate School of Medicine, Hokkaido University, Sapporo, 060–8638, Japan

Hiroshi Wakao

Contributed equally to this work with: Chie Sugimoto, Hiroshi Wakao

hwakao@med.hokudai.ac.jp

Affiliation Department of Hygiene & Cellular Preventive Medicine, Graduate School of Medicine, Hokkaido University, Sapporo, 060–8638, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK Hiroko Fujita Hiroyoshi Fujita HW. Performed the experiments: CS TK HW. Analyzed the data: CS RW TK Hiroko Fujita Hiroyoshi Fujita HW. Contributed reagents/materials/analysis tools: RW CS TK Hiroko Fujita Hiroyoshi Fujita HW. Wrote the paper: CS Hiroyoshi Fujita HW.