About the Authors

David L. Miller

dave@ninepointeightone.net

Affiliation Centre for Research into Ecological and Environmental Modelling, and School of Mathematics and Statistics, University of St Andrews, St Andrews, Scotland, United Kingdom

Len Thomas

Affiliation Centre for Research into Ecological and Environmental Modelling, and School of Mathematics and Statistics, University of St Andrews, St Andrews, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DLM LT. Performed the experiments: DLM. Analyzed the data: DLM. Contributed reagents/materials/analysis tools: DLM. Wrote the paper: DLM LT. Designed the software used in analysis: DLM LT.