About the Authors

Mark P. Dagleish

mark.dagleish@moredun.ac.uk

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Stuart Martin

Affiliation Animal Health & Veterinary Laboratories Agency Lasswade, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Philip Steele

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Jeanie Finlayson

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Samantha L. Eaton

Affiliation Neurobiology Division, The Roslin Institute at, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush Campus, Midlothian, EH25 9RG, United Kingdom

Sílvia Sisó

Affiliation Animal Health & Veterinary Laboratories Agency Lasswade, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Paula Stewart

Affiliation Neurobiology Division, The Roslin Institute at, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush Campus, Midlothian, EH25 9RG, United Kingdom

Natalia Fernández-Borges

Affiliation CIC bioGUNE, Parque tecnológico de Bizkaia, Derio 48160, Spain

Scott Hamilton

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Yvonne Pang

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Francesca Chianini

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Hugh W. Reid

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Wilfred Goldmann

Affiliation Neurobiology Division, The Roslin Institute at, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush Campus, Midlothian, EH25 9RG, United Kingdom

Lorenzo González

Affiliation Animal Health & Veterinary Laboratories Agency Lasswade, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Joaquín Castilla

Affiliations CIC bioGUNE, Parque tecnológico de Bizkaia, Derio 48160, Spain, IKERBASQUE, Basque Foundation for Science, Bilbao 48013, Bizkaia, Spain

Martin Jeffrey

Affiliation Animal Health & Veterinary Laboratories Agency Lasswade, Pentlands Science Park, Bush Loan, Penicuik, Near Edinburgh EH26 0PZ, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MJ MPD SM HWR JC. Performed the experiments: HWR MJ SM MPD LG SS P. Steele JF SH YP SLE FC NF-B WG P. Stewart JC. Analyzed the data: HWR MJ SM MPD LG SS P. Steele JF SH YP SLE FC NF-B WG P. Stewart JC. Contributed reagents/materials/analysis tools: HWR MJ SM MPD LG SS P. Steele JF SH YP SLE FC NF-B WG P. Stewart JC. Wrote the paper: HWR MJ SM MPD LG SS P. Steele JF SH YP SLE FC NF-B WG P. Stewart JC.