About the Authors

Tanja Opriessnig

Tanja.Opriessnig@roslin.ed.ac.uk

Affiliations The Roslin Institute and The Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, Midlothian, United Kingdom, Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Chao-Ting Xiao

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Priscilla F. Gerber

Affiliation The Roslin Institute and The Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, Midlothian, United Kingdom

Jianqiang Zhang

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Patrick G. Halbur

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors acknowledge that partial funding for this study was received from commercial sources (APC, Inc., Ankeny, Iowa, United States of America and EPFX/EGCO Protein, Provanco Feeds, Le Center, Minnesota, United States of America) and confirm that this funding did not interfere with the complete and objective presentation of data and this does not alter the author's adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TO PGH. Performed the experiments: TO PFG JZ CTX PGH. Analyzed the data: TO. Contributed reagents/materials/analysis tools: JZ. Contributed to the writing of the manuscript: TO PFG JZ CTX PGH.