About the Authors

Wenfa Xiao

Contributed equally to this work with: Wenfa Xiao, Xiaogai Ge

xiaowenf@caf.ac.cn (WX); gexiaogai2006@163.com (XG)

Affiliation State Forestry Administration Key Laboratory of Forest Ecology and Environment, Research Institute of Forest Ecology, Environment and Protection, Chinese Academy of Forestry, Beijing, PR China

Xiaogai Ge

Contributed equally to this work with: Wenfa Xiao, Xiaogai Ge

xiaowenf@caf.ac.cn (WX); gexiaogai2006@163.com (XG)

Affiliations State Forestry Administration Key Laboratory of Forest Ecology and Environment, Research Institute of Forest Ecology, Environment and Protection, Chinese Academy of Forestry, Beijing, PR China, Research Institute of Subtropical Forestry, Chinese Academy of Forestry, Fuyang, Zhejiang, PR China

Lixiong Zeng

Affiliation State Forestry Administration Key Laboratory of Forest Ecology and Environment, Research Institute of Forest Ecology, Environment and Protection, Chinese Academy of Forestry, Beijing, PR China

Zhilin Huang

Affiliation State Forestry Administration Key Laboratory of Forest Ecology and Environment, Research Institute of Forest Ecology, Environment and Protection, Chinese Academy of Forestry, Beijing, PR China

Jingpin Lei

Affiliation State Forestry Administration Key Laboratory of Forest Silviculture, Research Institute of Forestry, Chinese Academy of Forestry, Beijing, PR China

Benzhi Zhou

Affiliation Research Institute of Subtropical Forestry, Chinese Academy of Forestry, Fuyang, Zhejiang, PR China

Maihe Li

Affiliations Swiss Federal Research Institute WSL, Birmensdorf, Switzerland, State Key Laboratory of Forest and Soil Ecology, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, PR China

Competing Interests

This study was partly supported by CFERN&GENE Award Funds. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: WX XG. Performed the experiments: XG LZ ZH JL. Analyzed the data: XG ML. Contributed reagents/materials/analysis tools: XG WX ML. Wrote the paper: XG ML BZ.