About the Authors

Cristina Olivieri

Affiliation Laboratory of Molecular Archaeo-Anthropology/ancient DNA, School of Biosciences and Veterinary Medicine, University of Camerino, Camerino, Italy

Isolina Marota

Affiliation Laboratory of Molecular Archaeo-Anthropology/ancient DNA, School of Biosciences and Veterinary Medicine, University of Camerino, Camerino, Italy

Ermanno Rizzi

Affiliation Institute of Biomedical Technologies, National Research Council, Segrate, Italy

Luca Ermini

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Letizia Fusco

Affiliation Laboratory of Molecular Archaeo-Anthropology/ancient DNA, School of Biosciences and Veterinary Medicine, University of Camerino, Camerino, Italy

Alessandro Pietrelli

Affiliation Institute of Biomedical Technologies, National Research Council, Segrate, Italy

Gianluca De Bellis

Affiliation Institute of Biomedical Technologies, National Research Council, Segrate, Italy

Franco Rollo

Affiliation Laboratory of Molecular Archaeo-Anthropology/ancient DNA, School of Biosciences and Veterinary Medicine, University of Camerino, Camerino, Italy

Stefania Luciani

stefania.luciani@unicam.it

Affiliation Laboratory of Molecular Archaeo-Anthropology/ancient DNA, School of Biosciences and Veterinary Medicine, University of Camerino, Camerino, Italy

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CO SL GDB. Performed the experiments: CO LF IM ER. Analyzed the data: CO LE ER AP GDB LF FR. Wrote the paper: CO LE SL.