About the Authors
- Sayoko Ueda
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* E-mail: sykueda.ookami@gmail.com
Affiliations Tokyo Institute of Technology, Tokyo, Japan, Wildlife Research Center of Kyoto University, Kyoto, Japan
- Gaku Kumagai
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Affiliation Tama Zoological Park, Tokyo Zoological Park Society, Tokyo, Japan
- Yusuke Otaki
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Current address: Nogeyama Zoological Gardens, Yokohama, Japan
Affiliation Yokohama Zoological Gardens, Yokohama, Japan
- Shinya Yamaguchi
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Current address: Nogeyama Zoological Gardens, Yokohama, Japan
Affiliation Yokohama Zoological Gardens, Yokohama, Japan
- Shiro Kohshima
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Affiliation Wildlife Research Center of Kyoto University, Kyoto, Japan
Competing Interests
The authors have declared that no competing interests exist.
Author Contributions
Conceived and designed the experiments: SU SK. Performed the experiments: SU GK YO SY SK. Analyzed the data: SU SK. Contributed reagents/materials/analysis tools: GK YO SY. Wrote the paper: SU SK.