About the Authors

Thomas Pak

Contributed equally to this work with: Thomas Pak, Sooyeon Yoo, Ana M. Miranda-Angulo

Affiliation Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Sooyeon Yoo

Contributed equally to this work with: Thomas Pak, Sooyeon Yoo, Ana M. Miranda-Angulo

Affiliation Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Ana M. Miranda-Angulo

Contributed equally to this work with: Thomas Pak, Sooyeon Yoo, Ana M. Miranda-Angulo

Affiliations Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Institute of Medical Research, School of Medicine, Universidad de Antioquia, Medellín, Colombia

Hong Wang

Affiliation Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Seth Blackshaw

sblack@jhmi.edu

Affiliations Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Department of Ophthalmology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Department of Molecular Biology and Genetics, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Department of Neurology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Center for High-Throughput Biology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Institute for Cell Engineering, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: ALMA TP SY SB. Wrote the paper: ALMA TP SY SB. Conceived the project: ALMA SB. Conducted tamoxifen inductions and performed all histological experiments: TP SY. Bred and genotyped mice: HW. Designed the targeting vector and identified correctly targeted ES cells: ALMA.