About the Authors

Jacqueline T. Davis

Affiliation School of Mathematical Sciences, Queensland University of Technology, Brisbane, Australia

Kerrie Mengersen

Affiliation School of Mathematical Sciences, Queensland University of Technology, Brisbane, Australia

Nicola K. Abram

Affiliations Durrell Institute for Conservation and Ecology, School of Anthropology and Conservation, University of Kent, Canterbury, Kent, United Kingdom, HUTAN –Kinabatangan Orang-utan Conservation Programme, Sandakan, Sabah, Malaysia, Borneo Futures Project, People and Nature Consulting International, Jakarta, Indonesia

Marc Ancrenaz

Affiliations HUTAN –Kinabatangan Orang-utan Conservation Programme, Sandakan, Sabah, Malaysia, Borneo Futures Project, People and Nature Consulting International, Jakarta, Indonesia

Jessie A. Wells

Affiliation ARC Centre of Excellence for Environmental Decisions, University of Queensland, Brisbane, Australia

Erik Meijaard

emeijaard@gmail.com

Affiliations Borneo Futures Project, People and Nature Consulting International, Jakarta, Indonesia, ARC Centre of Excellence for Environmental Decisions, University of Queensland, Brisbane, Australia, School of Archaeology & Anthropology, Australian National University, Canberra, Australia

Competing Interests

Erik Meijaard is presently employed by People and Nature Consulting International (PNCI), but was not working for them at the time the study was conducted. Marc Ancrenaz and Nicola K. Abram are members of the Borneo Futures Project at PNCI. The work that went into analyzing and writing up the results of this study was unpaid. PNCI had no role in the study. The affiliation with PNCI does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KM EM. Performed the experiments: KM NKA JAW EM. Analyzed the data: JTD KM. Contributed reagents/materials/analysis tools: KM JAW EM. Wrote the paper: JTD KM NKA MA JAW EM.