About the Authors

David A. W. Miller

dxm84@psu.edu

Affiliations United States Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America, Pennsylvania State University, Department of Ecosystem Science and Management, University Park, Pennsylvania, United States of America

James D. Nichols

Affiliation United States Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Justin A. Gude

Affiliation Montana Fish, Wildlife and Parks, Helena, Montana, United States of America

Lindsey N. Rich

Affiliation United States Geological Survey, Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, Montana, United States of America

Kevin M. Podruzny

Affiliation Montana Fish, Wildlife and Parks, Helena, Montana, United States of America

James E. Hines

Affiliation United States Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Michael S. Mitchell

Affiliation United States Geological Survey, Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, Montana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JAG LNR KMP MSM. Performed the experiments: DAWM JDN JAG LNR KMP JEH MSM. Analyzed the data: DAWM JDN LNR JEH. Contributed reagents/materials/analysis tools: DAWM JEH. Wrote the paper: DAWM JDN JAG LNR KMP MSM.