About the Authors

Tara C. Smith

tara-smith@uiowa.edu

Affiliation Center for Emerging Infectious Diseases, Department of Epidemiology, University of Iowa College of Public Health, University of Iowa, Iowa City, Iowa, United States of America

Wondwossen A. Gebreyes

Affiliation Department of Veterinary Preventive Medicine, Ohio State University, Columbus, Ohio, United States of America

Melanie J. Abley

Affiliation Department of Veterinary Preventive Medicine, Ohio State University, Columbus, Ohio, United States of America

Abby L. Harper

Affiliation Center for Emerging Infectious Diseases, Department of Epidemiology, University of Iowa College of Public Health, University of Iowa, Iowa City, Iowa, United States of America

Brett M. Forshey

Affiliation Center for Emerging Infectious Diseases, Department of Epidemiology, University of Iowa College of Public Health, University of Iowa, Iowa City, Iowa, United States of America

Michael J. Male

Affiliation Center for Emerging Infectious Diseases, Department of Epidemiology, University of Iowa College of Public Health, University of Iowa, Iowa City, Iowa, United States of America

H. Wayne Martin

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Bayleyegn Z. Molla

Affiliation Department of Veterinary Preventive Medicine, Ohio State University, Columbus, Ohio, United States of America

Srinand Sreevatsan

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Siddhartha Thakur

Affiliation Department of Population Health and Pathobiology, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Madhumathi Thiruvengadam

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Peter R. Davies

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Competing Interests

TS is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: WG PD TS. Performed the experiments: TS MA AH MM HWM BMF SS ST MT. Analyzed the data: TS WG SS BZ BF. Wrote the paper: TS WG MA AH BF MM HWM BMF SS ST MT PD.