About the Authors

Zanna Clay

zannaclay@emory.edu

Affiliation Living Links, Yerkes National Primate Research Center, Emory University, Atlanta, Georgia, United States of America

Frans B. M. de Waal

Affiliation Living Links, Yerkes National Primate Research Center, Emory University, Atlanta, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZC FDW. Performed the experiments: ZC. Analyzed the data: ZC. Contributed reagents/materials/analysis tools: FDW. Wrote the paper: ZC FDW.