About the Authors

Brady J. Mattsson

brady.mattsson@gmail.com

Current address: Department of Forest- and Soil Sciences, University of Natural Resources and Life Sciences, Vienna, Austria

Affiliation Western Ecological Research Center, United States Geological Survey, Sacramento, California, United States of America

Elise F. Zipkin

Affiliation Patuxent Wildlife Research Center, United States Geological Survey, Laurel, Maryland, United States of America

Beth Gardner

Affiliation Department of Forestry and Environmental Resources, North Carolina State University, Raleigh, North Carolina, United States of America

Peter J. Blank

Current address: Department of Zoology. University of Wisconsin, Madison, Wisconsin, United States of America

Affiliation Patuxent Wildlife Research Center, United States Geological Survey, Laurel, Maryland, United States of America

John R. Sauer

Affiliation Patuxent Wildlife Research Center, United States Geological Survey, Laurel, Maryland, United States of America

J. Andrew Royle

Affiliation Patuxent Wildlife Research Center, United States Geological Survey, Laurel, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BJM EFZ JAR. Performed the experiments: BJM BG. Analyzed the data: BJM. Contributed reagents/materials/analysis tools: BG JAR PJB BG. Wrote the paper: BJM EFZ PJB JRS.