About the Authors

Jing Lv

Contributed equally to this work with: Jing Lv, Jianjian Qi, Qiuxiang Shi, Di Shen, Shengping Zhang

Current address: Tobacco Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Tobacco Biology and Processing, Ministry of Agriculture, Qingdao, China

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Jianjian Qi

Contributed equally to this work with: Jing Lv, Jianjian Qi, Qiuxiang Shi, Di Shen, Shengping Zhang

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Qiuxiang Shi

Contributed equally to this work with: Jing Lv, Jianjian Qi, Qiuxiang Shi, Di Shen, Shengping Zhang

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Di Shen

Contributed equally to this work with: Jing Lv, Jianjian Qi, Qiuxiang Shi, Di Shen, Shengping Zhang

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Shengping Zhang

Contributed equally to this work with: Jing Lv, Jianjian Qi, Qiuxiang Shi, Di Shen, Shengping Zhang

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Guangjin Shao

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Hang Li

Affiliations Middlebury College, Middlebury, Vermont, United States of America, High School Affiliated to Renmin University of China, Beijing, China

Zhanyong Sun

Affiliation East-West Seed International Ltd., Nanning, Guangxi, China

Yiqun Weng

Affiliation United States Department of Agriculture (USDA), ARS, Vegetable Crops Research Unit, Department of Horticulture, University of Wisconsin, Madison, Wisconsin, United States of America

Yi Shang

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Xingfang Gu

Current address: State Key Lab Crop Stress Biol Arid Areas, Coll Plant Protect, Northwest A&F University, Yangling, Shaanxi, China

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Xixiang Li

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Xiaoguo Zhu

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Jinzhe Zhang

Current address: National Laboratory for Protein Engineering and Plant Genetic Engineering, Peking-Yale Joint Research Center for Plant Molecular Genetics and AgroBiotechnology, College of Life Sciences, Peking University, Beijing, China

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Robbert van Treuren

Affiliation Centre for Genetic Resources, The Netherlands, Wageningen University and Research Centre, Wageningen, The Netherlands

Willem van Dooijeweert

Affiliation Centre for Genetic Resources, The Netherlands, Wageningen University and Research Centre, Wageningen, The Netherlands

Zhonghua Zhang

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Sanwen Huang

huangsanwen@caas.net.cn

Affiliation Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Sino-Dutch Joint Lab of Horticultural Genomics, Opening Lab of Genetic Improvement of Agricultural Crops of Ministry of Agriculture, Beijing, China

Competing Interests

Jing Lv is now at the Tobacco Research Institute of Chinese Academy of Agricultural Sciences/Key Laboratory of Tobacco Biology and Processing, Ministry of Agriculture, Qingdao, and thus receives tobacco funding. However this work was carried out at the Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, thus the research was not funded by TRI Zhanyong Sun is an employee of East-west Seed International, but this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials. There are no patents or marketed products related to this work.

Author Contributions

Conceived and designed the experiments: SH. Performed the experiments: QS JQ DS SZ GS HL JL YS XZ JZ. Analyzed the data: JL JQ QS ZZ RT WD ZS YW XG XL. Contributed reagents/materials/analysis tools: DS SZ ZS YW. Wrote the paper: JL JQ ZZ SH.