About the Authors

Scott A. Bernhardt

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Mark P. Simmons

Affiliation Department of Biology, Colorado State University, Fort Collins, Colorado, United States of America

Ken E. Olson

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Barry J. Beaty

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Carol D. Blair

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

William C. Black

wcb4@lamar.colostate.edu

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SAB CDB KEO BJB WCB. Performed the experiments: SAB MPS WCB. Analyzed the data: SAB MPS WCB. Contributed reagents/materials/analysis tools: MPS CDB KEO BJB WCB. Wrote the paper: SAB MPS CDB KEO WCB.