About the Authors

James A. Lutz

jlutz@uw.edu

Affiliation College of the Environment, University of Washington, Seattle, Washington, United States of America

Andrew J. Larson

Affiliation Department of Forest Management, University of Montana, Missoula, Montana, United States of America

Mark E. Swanson

Affiliation School of the Environment, Washington State University, Pullman, Washington, United States of America

James A. Freund

Affiliation School of Environmental and Forest Sciences, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JAL AJL MES JAF. Performed the experiments: JAL AJL MES JAF. Analyzed the data: JAL AJL MES JAF. Wrote the paper: JAL AJL MES.