About the Authors

Hua V. Lin

linhs@lilly.com

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Andrea Frassetto

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Edward J. Kowalik Jr

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Andrea R. Nawrocki

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Mofei M. Lu

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Jennifer R. Kosinski

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

James A. Hubert

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Daphne Szeto

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Xiaorui Yao

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Gail Forrest

Affiliation Diabetes and In Vivo Pharmacology, Merck Research Laboratories, Rahway, New Jersey, United States of America

Donald J. Marsh

Affiliation Safety Assessment, Merck Research Laboratories, West Point, Pennsylvania, United States of America

Competing Interests

All authors are employees of Merck Research Laboratories, Rahway. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HVL AF ARN DJM. Performed the experiments: HVL AF EJK ARN MML JRK JAH DS XY. Analyzed the data: HVL AF EJK DS XY GF. Wrote the paper: HVL DJM.