About the Authors

Mark A. McElwain

Affiliation Howard Hughes Medical Institute, Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Dennis C. Ko

Current address: Department of Microbiology, University of Washington, Seattle, Washington, United States of America

Affiliation Howard Hughes Medical Institute, Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Michael D. Gordon

Current address: Department of Zoology, University of British Columbia, Vancouver, British Columbia, Canada

Affiliation Howard Hughes Medical Institute, Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Henrik Fyrst

Affiliation Center for Cancer Research, Children's Hospital Oakland Research Institute, Oakland, California, United States of America

Julie D. Saba

Affiliation Center for Cancer Research, Children's Hospital Oakland Research Institute, Oakland, California, United States of America

Roel Nusse

rnusse@stanford.edu

Affiliation Howard Hughes Medical Institute, Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MAM DCK MDG HF JDS RN. Performed the experiments: MAM DCK MDG HF. Analyzed the data: MAM DCK MDG HF JDS RN. Contributed reagents/materials/analysis tools: MAM DCK MDG HF JDS RN. Wrote the paper: MAM HF.