About the Authors

Chunwang Li

Contributed equally to this work with: Chunwang Li, Xiaobo Yang

Affiliation Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Xiaobo Yang

Contributed equally to this work with: Chunwang Li, Xiaobo Yang

Affiliations Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, Graduate University of the Chinese Academy of Sciences, Beijing, China

Yuhua Ding

Affiliation Dafeng Milu National Nature Reserve, Dafeng Jiangsu, China

Linyuan Zhang

Affiliation Beijing Milu Ecological Research Center, Beijing, China

Hongxia Fang

Affiliation Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Songhua Tang

Affiliation Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Zhigang Jiang

jiangzg@ioz.ac.cn

Affiliations Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, Graduate University of the Chinese Academy of Sciences, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZJ CL. Performed the experiments: XY ZJ HF YD LZ. Analyzed the data: CL XY. Contributed reagents/materials/analysis tools: HF ST. Wrote the paper: CL ZJ XY.