About the Authors

Keyan Zhao

Affiliations Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America, Department of Genetics, Stanford University, Stanford, California, United States of America

Mark Wright

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Jennifer Kimball

Current address: Department of Crop Science, North Carolina State University, Raleigh, North Carolina, United States of America

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Georgia Eizenga

Affiliation Dale Bumpers National Rice Research Center, Agricultural Research Service (ARS), United States Department of Agriculture (USDA), Stuttgart, Arkansas, United States of America

Anna McClung

Affiliation Dale Bumpers National Rice Research Center, Agricultural Research Service (ARS), United States Department of Agriculture (USDA), Stuttgart, Arkansas, United States of America

Michael Kovach

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Wricha Tyagi

Current address: School of Crop Improvement, College of Post Graduate Studies, Central Agricultural University, Umiam, Meghalaya, India

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Md. Liakat Ali

Affiliation Rice Research and Extension Center, University of Arkansas, Stuttgart, Arkansas, United States of America

Chih-Wei Tung

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Andy Reynolds

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Carlos D. Bustamante

cdbustam@stanford.edu (CDB); srm4@cornell.edu (SRM)

Affiliations Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America, Department of Genetics, Stanford University, Stanford, California, United States of America

Susan R. McCouch

cdbustam@stanford.edu (CDB); srm4@cornell.edu (SRM)

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SRM KZ GE CDB. Performed the experiments: JK GE WT MLA CWT. Analyzed the data: KZ MHW AM MJK CWT AR. Wrote the paper: SRM KZ MHW CDB.